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Cirrate shell : ウィキペディア英語版
Cirrate shell

Cirrate octopuses possess a well-developed internal shell that supports their muscular swimming fins. This is in contrast to the more familiar, finless, incirrate octopuses, in which the shell remnant is either present as a pair of stylets or absent altogether.〔Fuchs, D., C. Ifrim & W. Stinnesbeck (2008). A new ''Palaeoctopus'' (Cephalopoda: Coleoidea) from the Late Cretaceous of Vallecillo, north-eastern Mexico, and implications for the evolution of Octopoda. ''Palaeontology'' 51(5): 1129–1139. 〕
The cirrate shell is quite unlike that of any other living cephalopod group and has its own dedicated set of descriptive terms.〔Young, R.E., M. Vecchione & K.M. Mangold (1999). (Cephalopoda Glossary ). Tree of Life Web Project.〕〔Young, R.E., M. Vecchione & K.M. Mangold (2002). (Cirrate Shell Terminology ). Tree of Life Web Project.〕 It is usually roughly arch- or saddle-shaped and is rather soft, being similar in consistency to cartilage.〔Vecchione, M., K.M. Mangold & R.E. Young (2010). (Cirrata Grimpe, 1916 ). Tree of Life Web Project.〕 Each of the eight extant cirrate genera is characterised by a distinct shell morphology:〔Collins, M.A. & R. Villanueva (2006). (Taxonomy, ecology and behaviour of the cirrate octopods ). In: Gibson, R.N., R.J.A. Atkinson & J.D.M. Gordon (eds.) ''Oceanography and Marine Biology: An Annual Review''. Aberdeen University Press, Aberdeen. pp. 277–322.〕
*Cirroteuthidae
*
*''Cirroteuthis'' — saddle-shaped, with large wings
*
*''Cirrothauma'' — butterfly-shaped
*Opisthoteuthidae
*
*''Cirroctopus'' — V-shaped, lateral wings tapering to fine points
*
*''Cryptoteuthis'' — U-shaped, each lateral wing ending in broad lobe with pointed projection〔Collins, M., R. Villanueva & R.E. Young (2008). (''Cryptoteuthis'' Collins, 2004 ). Tree of Life Web Project.〕
*
*''Grimpoteuthis'' — U-shaped, lateral wings ending bluntly
*
*''Luteuthis'' — W-shaped
*
*''Opisthoteuthis'' — U-shaped, lateral wings usually tapering to fine points but termination complex in certain species〔Villanueva, R., R.E. Young & M. Vecchione (2008). (''Opisthoteuthis'' Verrill 1883 ). Tree of Life Web Project.〕
*Stauroteuthidae
*
*''Stauroteuthis'' — U-shaped
The comparatively simple shells of Opisthoteuthidae and Stauroteuthidae are thought to approximate the ancestral shape, with those of Cirroteuthidae being more derived.〔Bizikov, V.A. (2004). (Evolution of shell in Octopodiformes (Cephalopoda) ). In: Abstracts of the conference ''Mollusks of the Northeastern Asia and Northern Pacific: Biodiversity, Ecology, Biogeography and Faunal History''. October 4–6, 2004, Vladivostok, Russia. pp. 21–23. 〕 The shell of ''Cirroctopus'' appears transitional in form between those of incirrate octopuses and other cirrates, and resembles the reduced shell of the Late Cretaceous ''Palaeoctopus newboldi''.〔 The paired, rod-shaped stylets of incirrates are evolutionarily derived from the lateral wings and horns of the cirrate shell.〔
==References==


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